Four adorable penguins live in Madagascar's Central Park Zoo. They are brave and witty. In order to celebrate the birthday of the fourth rookie, the boss took everyone into the most stringent gold storage in the United States. However, he was caught by an automatic cheese vending machine with tentacles. Turns out there's Dave the Octopus hidden in the cheese machine. Dave was the most popular star at the zoo years ago, but no one has watched Dave since the penguins appeared. In the future, Dave can only go to the major zoos. So Dave decides to take revenge on the penguins. He invented Medusa Serum to turn penguins into monsters. Knowing about it, the four penguins team up with the mysterious North Wind Secret Service. After unremitting efforts and ingenuity, they finally defeated this self-righteous "bad guy" and saved all the penguins
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